Maize has become a staple food
in many parts of the world, with the total production of maize surpassing
that of wheat
. In addition to being consumed directly by humans (often in the form of masa
), maize is also used for corn ethanol
, animal feed
and other maize products
, such as corn starch
and corn syrup
The six major types of maize are dent corn
, flint corn
, pod corn
, flour corn
, and sweet corn
Sugar-rich varieties called sweet corn are usually grown for human consumption as kernels, while field corn
varieties are used for animal feed, various corn-based human food uses (including grinding into cornmeal
, pressing into corn oil
, and fermentation and distillation into alcoholic beverages like bourbon whiskey
), and as chemical feedstocks. Maize is also used in making ethanol
and other biofuels
Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain.
In 2014, total world production was 1.04 billion tonnes
. Maize is the most widely grown grain crop
throughout the Americas, with 361 million metric tons
grown in the United States alone in 2014. Genetically modified maize
made up 85% of the maize planted in the United States in 2009. Subsidies in the United States
help to account for the high level of cultivation of maize in the United States and the fact that the U.S. is the world's largest maize producer.
Maize is a cultigen
; human intervention is required for it to propagate. Whether or not the kernels fall off the cob on their own is a key piece of evidence used in archaeology to distinguish domesticated maize from its naturally-propagating teosinte
Genetic evidence can also be used to determine when various lineages split.
Most historians believe maize was domesticated in the Tehuacán Valley
Recent research in the early 21st century has modified this view somewhat; scholars now indicate the adjacent Balsas River
Valley of south-central Mexico as the center of domestication.
An influential 2002 study by Matsuoka et al
. has demonstrated that, rather than the multiple independent domestications model, all maize arose from a single domestication in southern Mexico about 9,000 years ago. The study also demonstrated that the oldest surviving maize types are those of the Mexican highlands. Later, maize spread from this region over the Americas along two major paths. This is consistent with a model based on the archaeological record suggesting that maize diversified in the highlands of Mexico before spreading to the lowlands.
Archaeologist Dolores Piperno has said:
A large corpus of data indicates that it [maize] was dispersed into lower Central America by 7600 BP [5600 BC] and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP [5000–4000 BC].
— Dolores Piperno, The Origins of Plant Cultivation and Domestication in the New World Tropics: Patterns, Process, and New Developments
Since then, even earlier dates have been published.
According to a genetic study by Embrapa
, corn cultivation was introduced in South America from Mexico, in two great waves: the first, more than 6000 years ago, spread through the Andes
. Evidence of cultivation in Peru has been found dating to about 6700 years ago.
The second wave, about 2000 years ago, through the lowlands of South America.
The earliest maize plants grew only small, 25-millimetre-long (1 in) corn cobs, and only one per plant. In Jackson Spielvogel's view, many centuries of artificial selection (rather than the current view that maize was exploited by interplanting with teosinte
) by the indigenous people of the Americas resulted in the development of maize plants capable of growing several cobs per plant, which were usually several centimetres/inches long each.
cultivated maize in numerous varieties throughout Mesoamerica
; they cooked, ground and processed it through nixtamalization
. It was believed that beginning about 2500 BC, the crop spread through much of the Americas.
Research of the 21st century has established even earlier dates. The region developed a trade network based on surplus and varieties of maize crops.
of south-central Chile
cultivated maize along with quinoa
times; however, potato was the staple food
of most Mapuches, "specially in the southern and coastal [Mapuche] territories where maize did not reach maturity".
Before the expansion of the Inca Empire
maize was traded and transported as far south as 40°19' S in Melinquina, Lácar Department
In that location maize remains were found inside pottery dated to 730 ±80 BP and 920 ±60 BP. Probably this maize was brought across the Andes from Chile.
The presence of maize in Guaitecas Archipelago
(43°55' S), the southernmost outpost of pre-Hispanic agriculture,
is reported by early Spanish explorers.
However the Spanish may have misidentified the plant.
After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders carried it back to Europe
and introduced it to other countries. Spanish settlers far preferred wheat bread to maize, cassava
, or potatoes. Maize flour could not be substituted for wheat for communion bread, since in Christian
belief only wheat could undergo transubstantiation
and be transformed into the body of Christ.
Some Spaniards worried that by eating indigenous foods, which they did not consider nutritious, they would weaken and risk turning into Indians. "In the view of Europeans, it was the food they ate, even more than the environment in which they lived, that gave Amerindians and Spaniards both their distinctive physical characteristics and their characteristic personalities."
Despite these worries, Spaniards did consume maize. Archeological evidence from Florida sites indicate they cultivated it as well.
Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere.
Many small male flowers make up the male inflorescence, called the tassel.
The word maize
derives from the Spanish form of the indigenous Taíno
word for the plant, mahiz
It is known by other names around the world.
The word "corn" outside the US, Canada, Australia, and New Zealand refers to any cereal
crop, its meaning understood to vary geographically to refer to the local staple
In the United States,
Australia, and New Zealand, corn
primarily means maize; this usage started as a shortening of "Indian corn".
"Indian corn" primarily means maize (the staple grain of indigenous Americans
), but can refer more specifically to multicolored "flint corn
" used for decoration.
In places outside the US, Canada, Australia, and New Zealand, corn
often refers to maize in culinary contexts. The narrower meaning is usually indicated by some additional word, as in sweet corn
, corn on the cob
, baby corn
, the puffed confection known as popcorn
and the breakfast cereal known as corn flakes
In Southern Africa, maize is commonly called mielie
) or mealie
words derived from the Portuguese word for maize, milho
is preferred in formal, scientific, and international usage because it refers specifically to this one grain, unlike corn
, which has a complex variety of meanings that vary by context and geographic region. Maize
is used by agricultural bodies and research institutes
such as the FAO
. National agricultural and industry associations often include the word maize
in their name even in English-speaking countries where the local, informal word is something other than maize
; for example, the Maize Association of Australia, the Indian Maize Development Association, the Kenya Maize Consortium and Maize Breeders Network, the National Maize Association of Nigeria, the Zimbabwe Seed Maize Association. However, in commodities trading, corn
consistently refers to maize and not other grains.
Structure and physiology
The maize plant is often 3 m (10 ft) in height,
though some natural strains can grow 13 m (43 ft).
The stem is commonly composed of 20 internodes
of 18 cm (7 in) length.
The leaves arise from the nodes, alternately on opposite sides on the stalk.
A leaf, which grows from each node, is generally 9 cm (31
in) in width and 120 cm (3 ft 11 in) in length.
Ears develop above a few of the leaves in the midsection of the plant, between the stem and leaf sheath, elongating by around 3 mm (1
in) per day, to a length of 18 cm (7 in)
with 60 cm (24 in) being the maximum alleged in the subspecies.
They are female inflorescences
, tightly enveloped by several layers of ear leaves commonly called husks. Certain varieties of maize have been bred to produce many additional developed ears. These are the source of the "baby corn
" used as a vegetable in Asian cuisine
The apex of the stem ends in the tassel, an inflorescence
of male flowers. When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce
and release pollen. Maize pollen is anemophilous
(dispersed by wind), and because of its large settling velocity, most pollen falls within a few meters of the tassel.
, called silks
, emerge from the whorl of husk leaves at the end of the ear. They are often pale yellow and 18 cm (7 in) in length, like tufts of hair in appearance. At the end of each is a carpel, which may develop into a "kernel" if fertilized by a pollen grain. The pericarp
of the fruit is fused with the seed coat referred to as "caryopsis
", typical of the grasses
, and the entire kernel is often referred to as the "seed
". The cob is close to a multiple fruit
in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas
, and adhere in regular rows around a white, pithy substance, which forms the ear. The maximum size of kernels is reputedly 2.5 cm (1 in).
An ear commonly holds 600 kernels. They are of various colors: blackish, bluish-gray
, green, red, white and yellow. When ground into flour
, maize yields more flour with much less bran
than wheat does. It lacks the protein gluten
of wheat and, therefore, makes baked goods with poor rising capability. A genetic variant
that accumulates more sugar and less starch
in the ear is consumed as a vegetable and is called sweet corn
. Young ears can be consumed raw, with the cob
and silk, but as the plant matures (usually during the summer months), the cob becomes tougher and the silk dries to inedibility. By the end of the growing season
, the kernels dry out and become difficult to chew without cooking them tender first in boiling water.
Planting density affects multiple aspects of maize. Modern farming techniques in developed countries
usually rely on dense planting, which produces one ear per stalk.
Stands of silage
maize are yet denser,
and achieve a lower percentage of ears and more plant matter.
Maize is a facultative short-day plant
and flowers in a certain number of growing degree days
> 10 °C (50 °F) in the environment to which it is adapted.
The magnitude of the influence that long nights have on the number of days that must pass before maize flowers
is genetically prescribed
and regulated by the phytochrome
can be eccentric in tropical cultivars
such that the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels
Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one (DIMBOA
). DIMBOA is a member of a group of hydroxamic acids
(also known as benzoxazinoids) that serve as a natural defense against a wide range of pests, including insects, pathogenic
fungi and bacteria
. DIMBOA is also found in related grasses, particularly wheat. A maize mutant (bx) lacking DIMBOA is highly susceptible to attack by aphids
. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer
). As maize matures, DIMBOA levels and resistance to the corn borer decline.
Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.
Zea mays 'Ottofile giallo Tortonese'
Zea mays "strawberry"—MHNT
Zea mays "Oaxacan Green" MHNT
Variegated maize ears
Multicolored corn kernels (CSIRO
While yellow maizes derive their color from lutein
, in red-colored maizes, the kernel coloration is due to anthocyanins
. These latter substances are synthesized in the flavonoids synthetic pathway
from polymerization of flavan-4-ols
by the expression of maize pericarp color1 (p1) gene
which encodes an R2R3 myb
-like transcriptional activator
of the A1 gene encoding for the dihydroflavonol 4-reductase
while another gene (Suppressor of Pericarp Pigmentation 1 or SPP1) acts as a suppressor
The p1 gene encodes an Myb-homologous transcriptional activator of genes required for biosynthesis of red phlobaphene pigments, while the P1-wr allele specifies colorless kernel pericarp and red cobs, and unstable factor for orange1 (Ufo1) modifies P1-wr expression to confer pigmentation in kernel pericarp, as well as vegetative tissues, which normally do not accumulate significant amounts of phlobaphene pigments.
The maize P gene encodes a Myb homolog that recognizes the sequence CCT/AACC, in sharp contrast with the C/TAACGG bound by vertebrate Myb proteins.
Maize flowers may sometimes exhibit mutations that lead to the formation of female flowers in the tassel. These mutations, ts4
, prohibit the development of the stamen while simultaneously promoting pistil development.
This may cause inflorescences
containing both male and female flowers, or hermaphrodite
Exotic varieties of maize are collected to add genetic diversity
when selectively breeding new domestic strains
Maize is an annual grass in the family Gramineae
, which includes such plants as wheat
, and sugarcane
. There are two major species of the genus Zea
(out of six total): Zea mays
(maize) and Zea diploperennis
, which is a perennial
type of teosinte. The annual
teosinte variety called Zea mays mexicana
is the closest botanical relative to maize. It still grows in the wild as an annual in Mexico and Guatemala.
Many forms of maize are used for food, sometimes classified as various subspecies related to the amount of starch each has:
- Flour corn: Zea mays var. amylacea
- Popcorn: Zea mays var. everta
- Dent corn : Zea mays var. indentata
- Flint corn: Zea mays var. indurata
- Sweet corn: Zea mays var. saccharata and Zea mays var. rugosa
- Waxy corn: Zea mays var. ceratina
- Amylomaize: Zea mays
- Pod corn: Zea mays var. tunicata Larrañaga ex A. St. Hil.
- Striped maize: Zea mays var. japonica
This system has been replaced (though not entirely displaced) over the last 60 years by multivariable classifications based on ever more data. Agronomic
data were supplemented by botanical traits for a robust initial classification, then genetic, cytological
, protein and DNA evidence was added. Now, the categories are forms (little used), races, racial complexes, and recently branches.
The Maize Genetics Cooperation Stock Center, funded by the USDA Agricultural Research Service
and located in the Department of Crop Sciences at the University of Illinois at Urbana-Champaign
, is a stock center of maize mutants. The total collection has nearly 80,000 samples. The bulk of the collection consists of several hundred named genes, plus additional gene combinations and other heritable variants. There are about 1000 chromosomal aberrations (e.g., translocations and inversions) and stocks with abnormal chromosome numbers (e.g., tetraploids
). Genetic data describing the maize mutant stocks as well as myriad other data about maize genetics can be accessed at MaizeGDB
, the Maize Genetics and Genomics Database.
In 2005, the US National Science Foundation
(NSF), Department of Agriculture (USDA
) and the Department of Energy
(DOE) formed a consortium to sequence the B73 maize genome
. The resulting DNA sequence data was deposited immediately into GenBank
, a public repository for genome-sequence data. Sequences and genome annotations have also been made available throughout the project's lifetime at the project's official site.
Primary sequencing of the maize genome was completed in 2008.
On November 20, 2009, the consortium published results of its sequencing effort in Science
The genome, 85% of which is composed of transposons
, was found to contain 32,540 genes (By comparison, the human genome
contains about 2.9 billion bases and 26,000 genes). Much of the maize genome has been duplicated and reshuffled by helitrons
—group of rolling circle
Maize reproduces sexually each year. This randomly selects half the genes from a given plant to propagate to the next generation, meaning that desirable traits found in the crop (like high yield or good nutrition) can be lost in subsequent generations unless certain techniques are used.
Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of the earliest and most successful developing Reid's Yellow Dent in the 1860s. These early efforts were based on mass selection
. Later breeding efforts included ear to row selection (C. G. Hopkins c. 1896), hybrids made from selected inbred
lines (G. H. Shull, 1909), and the highly successful double cross hybrids using four inbred lines (D. F. Jones
c. 1918, 1922). University supported breeding programs were especially important in developing and introducing modern hybrids (Ref Jugenheimer Hybrid Maize Breeding and Seed Production pub. 1958). By the 1930s, companies such as Pioneer
devoted to production of hybrid maize had begun to influence long-term development. Internationally important seed banks such as the International Maize and Wheat Improvement Center
(CIMMYT) and the US bank at the Maize Genetics Cooperation Stock Center University of Illinois at Urbana-Champaign
important for future crop development.
Since the 1940s the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic modification have succeeded in increasing output and reducing the need for cropland, pesticides, water and fertilizer.
There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant.
Global maize program
CIMMYT operates a conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by the Drought Tolerant Maize for Africa project.
Genetically modified (GM) maize
was one of the 26 GM crops
grown commercially in 2016.
Grown since 1997 in the United States and Canada, 92% of the US maize crop was genetically modified in 2016
and 33% of the worldwide maize crop was GM in 2016.
As of 2011, Herbicide-tolerant maize varieties were grown in Argentina, Australia, Brazil, Canada, China, Colombia, El Salvador, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, the Russian Federation, Singapore, South Africa, Taiwan, Thailand, and the United States. Insect-resistant maize was grown in Argentina, Australia, Brazil, Canada, Chile, China, Colombia, Egypt, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, South Africa, Switzerland, Taiwan, the United States, and Uruguay.
In September 2000, up to $50 million worth of food products were recalled due to the presence of Starlink
genetically modified corn, which had been approved only for animal consumption and had not been approved for human consumption, and was subsequently withdrawn from the market.
Maize is the domesticated variant of teosinte
The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between the two is largely controlled by differences in just two genes, called grassy tillers-1 (gt1
) and teosinte branched-1 (tb1
Several theories had been proposed about the specific origin of maize in Mesoamerica:
- It is a direct domestication of a Mexican annual teosinte, Zea mays ssp. parviglumis, native to the Balsas River valley in south-eastern Mexico, with up to 12% of its genetic material obtained from Zea mays ssp. mexicana through introgression.
- It has been derived from hybridization between a small domesticated maize (a slightly changed form of a wild maize) and a teosinte of section Luxuriantes, either Z. luxurians or Z. diploperennis.
- It has undergone two or more domestications either of a wild maize or of a teosinte. (The term "teosinte" describes all species and subspecies in the genus Zea, excluding Zea mays ssp. mays.)
- It has evolved from a hybridization of Z. diploperennis by Tripsacum dactyloides.
In the late 1930s, Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum
, a related genus. This theory about the origin of maize has been refuted by modern genetic testing
, which refutes Mangelsdorf's model and the fourth listed above.:40
The teosinte origin theory was proposed by the Russian botanist Nikolai Ivanovich Vavilov
in 1931 and the later American Nobel Prize
-winner George Beadle
It is supported experimentally and by recent studies of the plants' genomes. Teosinte and maize can cross-breed and produce fertile offspring. A number of questions remain concerning the species, among them:
- how the immense diversity of the species of sect. Zea originated,
- how the tiny archaeological specimens of 3500–2700 BC could have been selected from a teosinte, and
- how domestication could have proceeded without leaving remains of teosinte or maize with teosintoid traits earlier than the earliest known until recently, dating from ca. 1100 BC.
of maize is of particular interest to researchers—archaeologists
, geographers, etc. The process is thought by some to have started 7,500 to 12,000 years ago. Research from the 1950s to 1970s originally focused on the hypothesis that maize domestication occurred in the highlands between the states of Oaxaca
, because the oldest archaeological remains of maize known at the time were found there.
Connection with 'parviglumis' subspecies teosinte
(top), maize-teosinte hybrid (middle), maize (bottom)
Genetic studies, published in 2004 by John Doebley
, identified Zea mays
, native to the Balsas River
valley in Mexico's southwestern highlands, and also known as Balsas teosinte, as being the crop wild relative
that is genetically most similar to modern maize.
This was confirmed by further studies, which refined this hypothesis somewhat. Archaeobotanical studies, published in 2009, point to the middle part of the Balsas River valley as the likely location of early domestication; this river is not very long, so these locations are not very distant. Stone milling tools with maize residue have been found in an 8,700 year old layer of deposits in a cave not far from Iguala, Guerrero
Doebley was part of the team that first published, in 2002, that maize had been domesticated only once, about 9,000 years ago, and then spread throughout the Americas.
A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at Guila Naquitz Cave
in the Oaxaca Valley
, date back roughly 6,250 years; the oldest ears from caves near Tehuacan
, Puebla, 5,450 B.P.
As maize was introduced to new cultures, new uses were developed and new varieties selected to better serve in those preparations. Maize was the staple food, or a major staple – along with squash
, Andean region potato
, and amaranth
– of most pre-Columbian
North American, Mesoamerican, South American, and Caribbean cultures. The Mesoamerican civilization, in particular, was deeply interrelated with maize. Its traditions and rituals involved all aspects of maize cultivation – from the planting to the food preparation. Maize formed the Mesoamerican people's identity.
It is unknown what precipitated its domestication, because the edible portion of the wild variety is too small, and hard to obtain, to be eaten directly, as each kernel is enclosed in a very hard bivalve shell.
In 1939, George Beadle demonstrated that the kernels of teosinte are readily "popped" for human consumption, like modern popcorn.
Some have argued it would have taken too many generations of selective breeding
to produce large, compressed ears for efficient cultivation. However, studies of the hybrids readily made by intercrossing teosinte and modern maize suggest this objection is not well founded.
Spreading to the north
Around 4,500 ago, maize began to spread to the north; it was first cultivated in what is now the United States at several sites in New Mexico and Arizona, about 4,100 ago.
During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for the new crop.
In 2005, research by the USDA Forest Service
suggested that the rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels
, which are very sensitive to environmental changes.
Seedlings three weeks after sowing
Because it is cold-intolerant, in the temperate zones
maize must be planted in the spring. Its root
system is generally shallow, so the plant is dependent on soil moisture. As a plant that uses C4 carbon fixation
, maize is a considerably more water-efficient crop than plants that use C3 carbon fixation
such as alfalfa
. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July
", although modern hybrids
generally exceed this growth rate. Maize used for silage
is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture is shown in many parts of Africa, where periodic drought
regularly causes maize crop failure and consequent famine
. Although it is grown mainly in wet, hot climates, it has been said to thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop.
Mature plants showing ears
Maize was planted by the Native Americans
in hills, in a complex system known to some as the Three Sisters
Maize provided support for beans
, and the beans provided nitrogen derived from nitrogen-fixing rhizobia
bacteria which live on the roots of beans and other legumes
; and squashes
provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil.
This method was replaced by single species hill planting where each hill 60–120 cm (2 ft 0 in–3 ft 11 in) apart was planted with three or four seeds, a method still used by home gardeners. A later technique was "checked maize", where hills were placed 1 m (40 in) apart in each direction, allowing cultivators to run through the field in two directions. In more arid lands, this was altered and seeds were planted in the bottom of 10–12 cm (4–41
in) deep furrows to collect water. Modern technique plants maize in rows which allows for cultivation while the plant is young, although the hill technique is still used in the maize fields of some Native American reservations. When maize is planted in rows, it also allows for planting of other crops between these rows to make more efficient use of land space.
In most regions today, maize grown in residential gardens
is still often planted manually with a hoe
, whereas maize grown commercially is no longer planted manually but rather is planted with a planter
. In North America, fields are often planted in a two-crop rotation
with a nitrogen-fixing
crop, often alfalfa
in cooler climates and soybeans
in regions with longer summers. Sometimes a third crop, winter wheat
, is added to the rotation.
Many of the maize varieties grown in the United States and Canada are hybrids. Often the varieties have been genetically modified
to tolerate glyphosate
or to provide protection against natural pests. Glyphosate is an herbicide which kills all plants except those with genetic tolerance. This genetic tolerance is very rarely found in nature.
In the midwestern United States, low-till or no-till farming
techniques are usually used. In low-till, fields are covered once, maybe twice, with a tillage implement either ahead of crop planting or after the previous harvest. The fields are planted and fertilized
are controlled through the use of herbicides
, and no cultivation tillage is done during the growing season. This technique reduces moisture evaporation from the soil, and thus provides more moisture for the crop. The technologies mentioned in the previous paragraph enable low-till and no-till farming. Weeds compete with the crop for moisture and nutrients, making them undesirable.
Mature maize ears
Hand-picking harvest of maize in Myanmar.
Before the 20th century, all maize harvesting was by manual labour
, by grazing
, or by some combination of those. Whether the ears were hand-picked and the stover
was grazed, or the whole plant was cut, gathered, and shocked
, people and livestock
did all the work. Between the 1890s and the 1970s, the technology of maize harvesting expanded greatly. Today, all such technologies, from entirely manual harvesting to entirely mechanized, are still in use to some degree, as appropriate to each farm's needs
, although the thoroughly mechanized versions predominate, as they offer the lowest unit costs
when scaled to large farm operations. For small farms, their unit cost can be too high, as their higher fixed cost
cannot be amortized
over as many units.
Before World War II
, most maize in North America was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees
). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover
) and corn binders, which are reaper-binders
designed specifically for maize (for example, Video
). The latter produce sheaves
that can be shocked
. By hand or mechanical picker, the entire ear is harvested, which then requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs
, and these whole ears are a sufficient form for some livestock feeding use. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine
and store it in bins
. The combine with a corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter
for the soil
. The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter the machinery. The combine separates the husk and the cob, keeping only the kernels.
When maize is a silage
crop, the entire plant is usually chopped at once with a forage harvester
(chopper) and ensiled in silos or polymer wrappers. Ensiling of sheaves cut by a corn binder was formerly common in some regions but has become uncommon.
Worldwide maize production
For storing grain in bins, the moisture of the grain must be sufficiently low to avoid spoiling. If the moisture content of the harvested grain is too high, grain dryers
are used to reduce the moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases (propane
or natural gas
) and electricity to power the blowers.
Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain.
In 2018, total world production was 1.15 billion tonnes
, led by the United States with 34.2% of the total (table). China produced 22.4% of the global total.
In 2016, maize production was forecast to be over 15 billion bushels
(380 million metric tons), an increase of 11% over 2014 American production.
Based on conditions as of August 2016, the expected yield would be the highest ever for the United States.
The area of harvested maize was forecast to be 87 million acres (352 000 km²), an increase of 7% over 2015.
Maize is especially popular in Midwestern
states such as Indiana
; in the latter, it was named the state's official grain in 2017.
The susceptibility of maize to the European corn borer and corn rootworms, and the resulting large crop losses which are estimated at a billion dollars worldwide for each pest,
led to the development of transgenics
expressing the Bacillus thuringiensis
toxin. "Bt maize" is widely grown in the United States and has been approved for release in Europe.
Poster showing a woman serving muffins, pancakes, and grits, with canisters on the table labeled corn meal, grits, and hominy, US Food Administration, 1918
In prehistoric times Mesoamerican
women used a metate
to process maize into ground cornmeal, allowing the preparation of foods that were more calorie dense than popcorn. After ceramic vessels were invented the Olmec
people began to cook maize together with beans, improving the nutritional value of the staple meal. Although maize naturally contains niacin
, an important nutrient, it was not bioavailable
without the process of nixtamalization
. The Maya
used nixtamal meal to make varieties of porridges and tamales.
The process was later used in the cuisine of the American South
to prepare corn for grits
Maize is a staple of Mexican cuisine
(cornmeal treated with limewater
) is the main ingredient for tortillas
and many other dishes of Central American food. It is the main ingredient of corn tortilla
and all the dishes based on them, like tacos
and many more. In Mexico the fungus of maize, known as huitlacoche
, is considered a delicacy.
made with corn meal
Boiled corn on a white plate
Maize can also be harvested and consumed in the unripe state, when the kernels are fully grown but still soft. Unripe maize must usually be cooked to become palatable; this may be done by simply boiling or roasting the whole ears and eating the kernels right off the cob. Sweet corn
, a genetic variety that is high in sugars and low in starch, is usually consumed in the unripe state. Such corn on the cob
is a common dish in the United States, Canada, United Kingdom, Cyprus, some parts of South America, and the Balkans, but virtually unheard of in some European countries.
Corn on the cob was hawked on the streets of early 19th-century New York City by poor, barefoot "Hot Corn
Girls", who were thus the precursors of hot dog carts
wagons, and fruit stands seen on the streets of big cities today.
Within the United States, the usage of maize for human consumption constitutes only around 1/40th of the amount grown in the country. In the United States and Canada, maize is mostly grown to feed livestock
, as forage, silage
(made by fermentation of chopped green cornstalks), or grain. Maize meal is also a significant ingredient of some commercial animal food products.
Sweetcorn, yellow, raw
(seeds only)Note: assuming freed niacin
Raw, yellow, sweet maize kernels are composed of 76% water, 19% carbohydrates
, 3% protein
, and 1% fat
(table). In a 100-gram
serving, maize kernels provide 86 calories
and are a good source (10–19% of the Daily Value
) of the B vitamins
(but see Pellagra warning
below), pantothenic acid
(B5) and folate
(right table for raw, uncooked kernels, USDA
Nutrient Database). In moderate amounts, they also supply dietary fiber
and the essential minerals
whereas other nutrients are in low amounts (table).
Feed and fodder for livestock
Maize is a major source of both grain feed
. It is fed to the livestock in various ways. When it is used as a grain crop, the dried kernels
are used as feed. They are often kept on the cob
for storage in a corn crib
, or they may be shelled off for storage in a grain bin
. The farm that consumes the feed may produce it, purchase it on the market, or some of both. When the grain is used for feed, the rest of the plant (the corn stover
) can be used later as fodder, bedding
(litter), or soil amendment
. When the whole maize plant (grain plus stalks and leaves) is used for fodder, it is usually chopped all at once
, as digestibility and palatability are higher in the ensilaged form than in the dried form. Maize silage is one of the most valuable forages for ruminants.
Before the advent of widespread ensilaging, it was traditional to gather the corn into shocks
after harvesting, where it dried further. With or without a subsequent move to the cover of a barn, it was then stored for weeks to several months until fed to the livestock. Today ensilaging can occur not only in siloes
but also in silage wrappers. However, in the tropics, maize can be harvested year-round and fed as green forage to the animals.
"Feed maize" is being used increasingly for heating;
specialized corn stoves
(similar to wood stoves
) are available and use either feed maize or wood pellets to generate heat. Maize cobs are also used as a biomass
fuel source. Maize is relatively cheap and home-heating furnaces have been developed which use maize kernels as a fuel. They feature a large hopper that feeds the uniformly sized maize kernels (or wood pellets or cherry
pits) into the fire.
Maize is increasingly used as a feedstock for the production of ethanol fuel
When considering where to construct an ethanol plant, one of the site selection criteria is to ensure there is locally available feedstock.
Ethanol is mixed with gasoline to decrease the amount of pollutants emitted when used to fuel motor vehicles. High fuel prices in mid-2007 led to higher demand for ethanol, which in turn led to higher prices paid to farmers for maize. This led to the 2007 harvest being one of the most profitable maize crops in modern history for farmers. Because of the relationship between fuel and maize, prices paid for the crop now tend to track the price of oil.
The price of food is affected to a certain degree by the use of maize for biofuel production. The cost of transportation, production, and marketing are a large portion (80%) of the price of food in the United States. Higher energy costs affect these costs, especially transportation. The increase in food prices
the consumer has been seeing is mainly due to the higher energy cost. The effect of biofuel production on other food crop prices is indirect. Use of maize for biofuel production increases the demand, and therefore price of maize. This, in turn, results in farm acreage being diverted from other food crops to maize production. This reduces the supply of the other food crops and increases their prices.
Farm-based maize silage digester located near Neumünster
in Germany, 2007. Green inflatable biogas holder is shown on top of the digester
Maize is widely used in Germany as a feedstock for biogas plants
. Here the maize is harvested, shredded then placed in silage
clamps from which it is fed into the biogas plants. This process makes use of the whole plant rather than simply using the kernels as in the production of fuel ethanol.
Increasingly, ethanol is being used at low concentrations (10% or less) as an additive in gasoline
) for motor fuels to increase the octane rating
, lower pollutants, and reduce petroleum use (what is nowadays also known as "biofuels
" and has been generating an intense debate regarding the human beings' necessity of new sources of energy, on the one hand, and the need to maintain, in regions such as Latin America, the food habits and culture which has been the essence of civilizations such as the one originated in Mesoamerica; the entry, January 2008, of maize among the commercial agreements of NAFTA
has increased this debate, considering the bad labor conditions of workers in the fields, and mainly the fact that NAFTA "opened the doors to the import of maize from the United States, where the farmers who grow it receive multimillion-dollar subsidies and other government supports. ... According to OXFAM UK, after NAFTA went into effect, the price of maize in Mexico fell 70% between 1994 and 2001. The number of farm jobs dropped as well: from 8.1 million in 1993 to 6.8 million in 2002. Many of those who found themselves without work were small-scale maize growers.").
However, introduction in the northern latitudes of the US of tropical maize for biofuels
, and not for human or animal consumption, may potentially alleviate this.
Ornamental and other uses
Some forms of the plant are occasionally grown for ornamental use in the garden. For this purpose, variegated and colored leaf forms as well as those with colorful ears are used.
Corncobs can be hollowed out and treated to make inexpensive smoking pipes
, first manufactured in the United States in 1869.
Children playing in a maize kernel box
An unusual use for maize is to create a "corn maze
" (or "maize maze") as a tourist attraction. The idea of a maize maze was introduced by the American Maze Company who created a maze in Pennsylvania
in 1993.[better source needed]
Traditional mazes are most commonly grown using yew hedges
, but these take several years to mature. The rapid growth of a field of maize allows a maze to be laid out using GPS
at the start of a growing season and for the maize to grow tall enough to obstruct a visitor's line of sight by the start of the summer. In Canada and the US, these are popular in many farming communities.
Maize kernels can be used in place of sand in a sandboxlike
enclosure for children's play.
Additionally, feed corn is sometimes used by hunters to bait animals such as deer or wild hogs.
United States usage breakdown
The breakdown of usage of the 12.1-billion-bushel
(307-million-tonne) 2008 US maize crop was as follows, according to the World Agricultural Supply and Demand Estimates Report by the USDA.
In the US since 2009/2010, maize feedstock use for ethanol production has somewhat exceeded direct use for livestock feed; maize use for fuel ethanol was 5,130 million bushels (130 million tonnes) in the 2013/2014 marketing year.
A fraction of the maize feedstock dry matter used for ethanol production is usefully recovered as DDGS (dried distillers grains with solubles). In the 2010/2011 marketing year, about 29.1 million tonnes of DDGS were fed to US livestock and poultry. Because starch utilization in fermentation for ethanol production leaves other grain constituents more concentrated in the residue, the feed value per kg of DDGS, with regard to ruminant-metabolizable energy and protein, exceeds that of the grain. Feed value for monogastric animals, such as swine and poultry, is somewhat lower than for ruminants.
Comparison to other staple foods
Nutrient contents in %DV of common foods (raw, uncooked) per 100 g
The following table shows the nutrient content of maize and major staple foods in a raw harvested form. Raw forms are not edible and cannot be digested. These must be sprouted, or prepared and cooked for human consumption. In sprouted or cooked form, the relative nutritional and anti-nutritional contents of each of these staples are different from that of raw form of these staples reported in the table below.
Note: niacin for maize assumes freed niacin. A
raw yellow dent cornB
raw unenriched long-grain white riceC
raw hard red winter wheatD
raw potato with flesh and skinE
raw green soybeansG
raw sweet potatoH
When maize was first introduced into farming systems other than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food
. This was a mystery, since these types of malnutrition were not normally seen among the indigenous Americans, for whom maize was the principal staple food.
It was eventually discovered that the indigenous Americans had learned to soak maize in alkali
-water (the process now known as nixtamalization
) —made with ashes and lime (calcium oxide
) since at least 1200–1500 BC by Mesoamericans
and North Americans—which liberates the B-vitamin niacin
, the lack of which was the underlying cause of the condition known as pellagra
Maize was introduced into the diet of non-indigenous Americans without the necessary cultural knowledge acquired over thousands of years in the Americas. In the late 19th century, pellagra reached epidemic proportions in parts of the southern US, as medical researchers debated two theories for its origin: the deficiency theory (which was eventually shown to be true) said that pellagra was due to a deficiency of some nutrient, and the germ theory said that pellagra was caused by a germ transmitted by stable flies. A third theory, promoted by the eugenicist Charles Davenport
, held that people only contracted pellagra if they were susceptible to it due to certain "constitutional, inheritable" traits of the affected individual.
Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high lysine maize and the promotion of a more balanced diet have also contributed to its demise. Pellagra still exists today in food-poor areas and refugee camps where people survive on donated maize.
Maize has been an essential crop in the Andes
since the pre-Columbian era
. The Moche
culture from Northern Peru made ceramics from earth, water, and fire. This pottery was a sacred substance, formed in significant shapes and used to represent important themes. Maize was represented anthropomorphically as well as naturally.
In the United States, maize ears along with tobacco leaves are carved into the capitals of columns in the United States Capitol
building. Maize itself is sometimes used for temporary architectural detailing when the intent is to celebrate the fall season, local agricultural productivity and culture. Bundles of dried maize stalks are often displayed along with pumpkins, gourds and straw in autumnal displays outside homes and businesses. A well-known example of architectural use is the Corn Palace
in Mitchell, South Dakota, which uses cobs and ears of colored maize to implement a mural design that is recycled annually. Another well-known example is the Field of Corn
sculpture in Dublin, Ohio
, where hundreds of concrete ears of corn stand in a grassy field.
A maize stalk with two ripe ears is depicted on the reverse
of the Croatian 1 lipa
coin, minted since 1993.
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